1. Department of Neuromuscular and Developmental Anatomy, Division of Bioregulatory Medicine, Gunma University Graduate School of Medicine
2. The 21st Century Center of Excellence Program for Biomedical Research Using Accelerator Technology
3. Microbeam Radiation Biology Group, Radiation-applied Biology Division, Quantum Beam Science Directorate, Japan Atomic Energy Agency
4. Department of Quantum Biology, Division of Bioregulatory Medicine, Gunma University Graduate School of Medicine
5. Present address: Laboratory Veterinary Radiology, School of Veterinary Medicine and Animal Science, Kitasato University