1. Division of Animal, Horticultural, and Food Sciences, Department of Horticulture, Chungbuk National University, Cheongju 28644, Korea
2. Brain Korea 21 Center for Bio-Resource Development, Chungbuk National University, Cheongju 28644, Korea
3. Division of Plant Resources, Korea National Arboretum, Yangpyeong 12519, Korea