1. Institut de Mathématiques de Toulouse, UMR5219, Université de Toulouse, CNRS, UPS, F-31062 Toulouse Cedex 9, France
2. Department of Mathematics, Imperial College London, South Kensington Campus, London, SW7 2AZ, UK
3. Institute for the Advanced Study of Human Biology (ASHBi), Kyoto University Institute for Advanced Study, Kyoto University, Kyoto 606-8315, Japan