1. State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, P. R. China
2. Graduate University of Chinese of Academy Sciences, Beijing, 100049, P. R. China
3. Research Group Plant and Vegetation Ecology, Department of Biology, University of Antwerp, (Campus Drie Eiken), Universiteitsplein 1, B-2610 Wilrijk, Belgium
4. College of Information Science and Engineering, Shandong University of Science and Technology, Qingdao 266510, P. R. China
5. Department of Ecology and Environmental Science, & Umeå Marine Sciences Centre, Umeå University, SE-901 87 Umeå, Swede