1. College of Forestry, Central South University of Forestry and Technology, 498 Shaoshan Nanlu, Changsha 410004, Hunan, P. R. China
2. Research Institute of Forest, Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, P. R. China
3. Center for Statistical Genetics, Pennsylvania State University, Loc T3436, Mailcode CH69, 500 University Drive, Hershey PA 17033, USA