1. Centre for Mathematical Medicine and Biology, School of Mathematical Sciences, University of Nottingham, University Park, Nottingham, NG7 2RD, UK
2. INRIA Paris-Rocquencourt Research Center, Domaine de Voluceau – BP 105, 78153 Le Chesnay Cedex, France
3. School of Computing and Mathematics, University of Plymouth, Portland Square A316, Plymouth, PL4 8AA, UK