1. Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Science, Qingdao 266071, Shandong Province, P. R. China
2. Joint Laboratory for Deep Blue Fishery Engineering, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao, 266237, Shandong province, P. R. China
3. Department of Marine Technology, Ocean University of China, Qingdao 266100, Shandong Province, P. R. China