1. Edward C. Dickinson, Flat 19, Marlborough Court, Southfields Road, Eastbourne, East Sussex BN21 1BT, UK; and Naturalis Biodiversity Center, Darwinweg 2, 2333 CR, Leiden, the Netherlands, e-mail: .
2. Pepijn Kamminga, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR, Leiden, the Netherlands, e-mail: .
3. Steven van der Mije, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR, Leiden, the Netherlands, e-mail: