1. Department of Biological and Environmental Sciences, University of West Alabama, Livingston, AL 35470 USA; and Edge Engineering and Science, Houston, TX 77084 USA
2. Fisheries Division, Kentucky Department of Fish and Wildlife Resources, Frankfort, KY 40601 USA
3. Fisheries and Aquatic Monitoring, Tennessee Valley Authority, Chattanooga, TN 37402 USA
4. Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520-8106 USA