1. School of Biological and Environmental Sciences, Liverpool John Moores University, Liverpool L3 3AF United Kingdom
2. Centro de Apoio à Pesquisa Paleontológica da Quarta Colônia, Universidade Federal de Santa Maria, São João do Polêsine RS 97230-000 Brazil
3. Department of Environmental Earth Science, Eastern Connecticut State University, Willimantic, CT 06226 USA