1. Department of Applied Ecology, North Carolina State University, Raleigh, NC 27695 USA
2. Senckenberg Biodiversity and Climate Research Institute, Frankfurt am Main 60325 Germany
3. Division of Vertebrate Zoology, Peabody Museum of Natural History, Yale University, New Haven, CT 06520 USA
4. College of Veterinary Medicine, North Carolina State University, Raleigh, NC 27607 USA