1. Forestry and Forest Products Research Institute, Matsunosato 1, Tsukuba, Ibaraki, 305–8687, Japan;
2. Tohoku Research Center, Forestry and Forest Products Research Institute, 92–25 Nabeyashiki, shimokuriyagawa, Morioka, Iwate 020–0123 Japan;
3. E‐mail
4. Hokkaido Research Center, Forestry and Forest Products Research Institute, Hitsujigaoka, Toyohira, Sapporo, Hokkaido 062–8516, Japan;
5. Faculty of Education, Shizuoka University, 836 Ohtani, Suruga, Shizuoka 422–8529 Japan;
6. Graduate School of Life Sciences, Tohoku University, 6–3 Aoba, Aramaki, Aoba‐ku, Sendai, 980–8578 Japan;