1. Smithsonian Tropical Research Institute, Apartado 0843–03092, Balboa, Ancón, Panama
2. Present address: Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544 USA
3. Biological Dynamics of Forest Fragments Project, National Institute for Amazon Research (INPA), C.P. 478, 69011–970 Manaus, AM, Brazil
4. E‐mail
5. Department of Ecology, National Institute for Amazon Research (INPA), C.P. 478, 69011–970, Manaus, AM, Brazil