1. Department of Entomology and Academic Programs, College of Agricultural, Human, and Natural Resource Sciences, Washington State University, Pullman, Washington 99164, U.S.A.
2. Systematic Entomology Laboratory, Plant Sciences Institute, ARS, USDA, c/o National Museum of Natural History, MRC-168, P.O. Box 37012, Washington, DC 20013-7012, U.S.A.
3. Centro de Estudios Ambientales y Biodiversidad, Universidad del Valle de Guatemala 18 Avenida 11-95 Zona 15, Vista Hermosa III, Guatemala, Guatemala, C.A.