1. Department of Biological Sciences, Simon Fraser University, Burnaby, British Columbia V5A 1S6, Canada
2. Agassiz Research and Development Centre, Agriculture and Agri-Food Canada, Agassiz, British Columbia V0M 1A0, Canada
3. Centre de recherche sur les grains, Saint-Mathieu-de-Beloeil, Quebec J3G 0E2, Canada
4. Ottawa Research and Development Centre, Agriculture and Agri-Food Canada, Ottawa, Ontario K1A 0C6, Canada