1. Hokkaido National Fisheries Research Institute, Japan Fisheries Research and Education Agency
2. Faculty of Science and Technology, Oita University
3. National Research Institute of Fisheries Science, Japan Fisheries Research and Education Agency
4. Shibetsu Salmon Museum
5. Fisheries Research Center, Toyama Prefectual Agricultural Forestry and Fisheries Research Center
6. Tochigi Prefectural Fisheries Experiment Station