1. Simons Center for Quantitative Biology, Cold Spring Harbor Laboratory, Cold Spring Harbor, NY, USA
2. Department of Natural Resource Management, South Dakota State University, Brookings, SD, USA
3. Department of Botany, National Museum of Natural History, MRC 166, Smithsonian Institution, Washington, DC, USA
4. Department of Biology and Microbiology, South Dakota State University, Brookings, SD, USA