1. Max Planck Institute for Molecular Genetics, Ihnestrasse 63–73 14195, Berlin, Germany
2. Department of Biostatistics, University of North Carolina at Chapel Hill, NC 27599, USA
3. Department of Cellular and Molecular Pharmacology, University of California, San Francisco, CA 94158, USA
4. Computational Systems Biology, Institut de Biologie de l'Ecole Normale Supérieure (IBENS), CNRS, Inserm, Ecole Normale Supérieure, PSL Research University, F-75005 Paris, France