1. Department of Botany, Faculty of Science, Kasetsart University, Bangkok, Thailand
2. Center for Advanced Studies in Tropical Natural Resources, NRU-KU, Kasetsart University, Chatuchak, Bangkok, Thailand
3. Science and Education, The Field Museum, Chicago, IL, USA
4. Department of Biology and M.L. Bean Life Science Museum, Brigham Young University, Provo, UT, USA