1. Department of Ecology & Evolutionary Biology, University of Tennessee , Knoxville TN 37996 , USA
2. Missouri Botanical Garden , St. Louis, MO 63121 , USA
3. UNC-CH Herbarium (NCU), North Carolina Botanical Garden, University of North Carolina at Chapel Hill , C.B. 3280, Chapel Hill NC 27599-3280 , USA
4. Biology Department, Washington & Lee University , Lexington VA 24450 , USA