1. Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AE, UK
2. Institute of Evolutionary Biology, University of Edinburgh, Ashworth Laboratories, Charlotte Auerbach Road, Edinburgh, EH9 3FL, UK
3. Royal Botanic Garden Edinburgh, 20a Inverleith Row, Edinburgh, EH3 5LR, UK
4. Geography, University of Exeter, Laver Building, North Park Road, Exeter, EX4 4QE, UK