1. Jardín Botánico Lankester, Universidad de Costa Rica, carretera Cartago-Paraíso, Cartago, Costa Rica
2. Herbarium UCH, Universidad Autónoma de Chiriquí, Ciudad Universitaria, El Cabrero, Vía Interamericana, David, Chiriquí, Panama
3. Naturalis Biodiversity Center, Endless Forms, Sylviusweg, The Netherlands
4. Institute Biology Leiden, Leiden University, Sylviusweg, The Netherlands
5. Royal British Columbia Museum and American Museum of Natural History, SE, Salmon Arm, Canada
6. Science and Technology, University of Applied Sciences Leiden, Zernikedreef, CK, The Netherlands
7. Department of Evolution and Ecology, University of California, Davis, CA, USA
8. KU Leuven, Ecology, Evolution and Biodiversity Conservation, Charles Deberiotstraat, Leuven, Belgium