1. School of Life Sciences, University of Sussex, Falmer, Brighton, Sussex, UK
2. Centre for Ecology, Evolution and Conservation, School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, UK
3. Royal Botanic Gardens, Kew, Surrey, UK
4. Durrell Institute of Conservation and Ecology, School of Anthropology and Conservation, Marlowe Building, University of Kent, Canterbury, Kent, UK