1. Department of Biological Sciences, College of Arts and Sciences, University of the East, Manila, Philippines
2. The Graduate School, University of Santo Tomas, Espana, Manila, Philippines
3. Department of Plant Systematics, University of Bayreuth, Universitätstraße, Bayreuth, Germany College of Science and Research Center for the Natural & Applied Sciences, University of Santo Tomas, España, Manila, Philippines
4. The Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, UK
5. College of Science and Research Center for the Natural & Applied Sciences, University of Santo Tomas, España, Manila, Philippines