1. Department of Biology, University of Hawai‘i at Hilo , 200 W Kāwili St, Hilo, Hawai‘i 96720 , United States
2. U.S. Geological Survey, Pacific Island Ecosystems Research Center , PO Box 44, Hawai‘i National Park, Hawai‘i 96718 , United States
3. Center for Conservation Genomics, National Zoo and Conservation Biology Institute, Smithsonian Institution , Washington DC 20008 , United States
4. San Diego Zoo Wildlife Alliance , P.O. Box 39, Volcano, HI 96785 , United States
5. Centre for Biological Diversity, School of Biology, University of St Andrews , St Andrews KY16 9TH , United Kingdom