1. chool of Biological Sciences, Tokai University Department of Marine Biology and Sciences, S , Sapporo, Hokkaido , Japan
2. Mariculture Fisheries Research Institute, Fisheries Research Department, Hokkaido Research Organization , Muroran, Hokkaido , Japan
3. Tropical Biosphere Research Center, University of the Ryukyus , Nishihara, Okinawa , Japan
4. Faculty of Agriculture, Ryukoku University , Otsu, Shiga , Japan
5. Institute for Advanced Biosciences, Keio University , Tsuruoka, Yamagata , Japan
6. Department of Applied Aquabiology, National Fisheries University , Shimonoseki, Yamaguchi , Japan