1. Department of Natural Sciences, Faculty of Education, University of the Ryukyus, Nishihara, Okinawa, Japan
2. Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu, Sakyo, Kyoto, Japan
3. Center for Molecular Biodiversity Research, National Museum of Nature and Science, Tsukuba, Ibaraki, Japan
4. Kitakyushu Museum of Natural History and Human History, Higashida, Yahatahigashiku, Kitakyushu, Fukuoka, Japan