1. School of Marine Sciences, University of Maine, Orono , ME 04469 , US
2. Bedford Institute of Oceanography , Dartmouth, NS B2Y 4A2 , Canada
3. Fisheries and Oceans Canada, Maurice Lamontagne Institute , Mont-Joli, QC G5H 3Z4 , Canada
4. Department of Infectious Disease and Global Health, Cummings School of Veterinary Medicine, Tufts University , North Grafton, MA 01536 , US
5. Protected Species Branch, NOAA, NMFS, Northeast Fisheries Science Center , Woods Hole, MA 02543 , USA