1. Department of Biomolecular Engineering, University of California–Santa Cruz , Santa Cruz, CA 95064 , USA
2. Department of Medicine, Knight Cardiovascular Institute, Oregon Health and Science University , Portland, OR 97239 , USA
3. University of California Santa Cruz Genomics Institute , Santa Cruz, CA 95064 , USA
4. Department of Biology, Pennsylvania State University , University Park, PA , USA
5. Department of Molecular and Cell Biology, University of Connecticut , Storrs, CT 06296 , USA
6. Institute for Systems Genomics, University of Connecticut , Storrs, CT 06296 , USA
7. Center for Medical Genomics, Pennsylvania State University , University Park, PA , USA
8. Center for Computational Biology and Bioinformatics, Pennsylvania State University , University Park, PA , USA
9. Department of Computer Science and Engineering, Pennsylvania State University , University Park, PA , USA
10. Department of Biochemistry and Molecular Biology, Pennsylvania State University , University Park, PA , USA
11. Department of Molecular and Medical Genetics, Oregon Health and Science University , Portland, OR 97239 , USA
12. Division of Genetics, Oregon National Primate Research Center , Beaverton, OR 97006 , USA
13. Department of Informatics and Clinical Epidemiology, Oregon Health and Science University , Portland, OR 97239 , USA