1. Graduate School of Natural Science and Technology, Okayama University, Kita-ku, Okayama, Okayama, Japan
2. Institute of Environmental Toxicology, Joso, Ibaraki, Japan
3. The Jackson Laboratory, Bar Harbor, ME, Japan
4. Graduate School of Environmental and Life Science, Okayama University, Kita-ku, Okayama, Okayama, Japan
5. Laboratory of Pathology and Development, Institute for Quantitative Biosciences, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan