1. Laboratory of Animal Physiology, Graduate School of Agricultural Science, Tohoku University, Aoba-ku, Sendai, Japan
2. Research Faculty of Agriculture, Hokkaido University, Kita-ku, Sapporo, Japan
3. Division of Grassland Farming, Institute of Livestock and Grassland Science, National Agriculture and Food Research Organization (NARO), Nasushiobara, Tochigi, Japan
4. Division of Dairy Production Research, Hokkaido Agricultural Research Center, National Agriculture and Food Research Organization (NARO), Toyohira-ku, Sapporo, Japan
5. Laboratory of Meat Science, Graduate School of Agriculture, National University Cooperation Kagoshima University, Korimoto, Kagoshima-shi, Japan
6. Department of Agricultural Biotechnology and Research Institute for Agriculture and Life Sciences, College of Agriculture and Life Sciences, Seoul National University, Seoul, Republic of Korea