1. Agriculture and Agri-Food Canada, London Research and Development Centre, London, Ontario N5V 4T3, Canada
2. Department of Biology, Western University, London, Ontario N6A 5B7, Canada
3. School of Life Sciences, Guangdong Provincial Key Laboratory of Plant Resource, Sun Yat-sen University, Guangzhou 510275, Guangdong, China
4. Agriculture and Agri-Food Canada, Harrow Research and Development Centre, Harrow, Ontario N0R 1G0, Canada
5. Guangdong Academy of Agricultural Sciences, Guangzhou 510640, China