1. Department of Biochemistry, Kyushu University Graduate School of Medical Sciences, 3-1-1 Maidashi, Higashi-ku, Fukuoka 812-8582, Japan
2. Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency (JST), 3-1-1 Maidashi, Higashi-ku, Fukuoka 812-8582, Japan