1. Department of Biological Sciences, Northern Arizona University , Flagstaff, AZ 86011 , USA
2. Department of Biology and Ecology Center, Utah State University , Moab, UT 84532 , USA
3. Division of Biological Sciences, University of Montana , Missoula, MT 59812 , USA
4. Department of Biological Sciences, California State University , Los Angeles, CA 90032 , USA
5. Northeast Ohio Medical University , Rootstown, OH 44272 , USA