1. Department of Life Sciences, School of Agriculture, Meiji University , 1-1-1, Higashimita, Tama-ku, Kawasaki, Kanagawa, 214-8571 Japan
2. School of Biological and Environmental Sciences, Kwansei Gakuin University , Gakuen Uegahara 1, Sanda, Hyogo, 669-1330 Japan