1. Division of Biological Science, Graduate School of Science and Technology, Nara Institute of Science and Technology , Ikoma, Nara, 630-0192 Japan
2. RIKEN Center for Sustainable Resource Science , Yokohama, Kanagawa, 230-0045 Japan
3. Department of Botany and Plant Sciences, University of California, Riverside , Riverside, CA 92507, USA
4. RIKEN BioResource Research Center , Tsukuba, Ibaraki, 305-0074 Japan
5. Graduate School of Science, University of Tokyo , Hongo, Tokyo, 113-0033 Japan