1. Graduate School of Science and Technology, Nara Institute of Science and Technology , 8916-5 Takayama, Ikoma, Nara, 630-0192 Japan
2. Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University , Kurokami 2-39-1, Chuo-ku, Kumamoto, 860-8555 Japan
3. International Research Organization for Advanced Science and Technology, Kumamoto University , Kurokami 2-39-1, Chuo-ku, Kumamoto, 860-8555 Japan
4. Division of Cellular Dynamics, National Institute for Basic Biology , Nishigonaka 38, Myodaiji, Okazaki, Aichi, 444-8585 Japan
5. Department of Basic Biology, SOKENDAI (The Graduate University for Advanced Studies) , Nishigonaka 38, Myodaiji, Okazaki, Aichi, 444-8585 Japan
6. Research Center for Advanced Science and Technology, The University of Tokyo , 4-6-1 Komaba, Tokyo, 153-8904 Japan