1. Biology Department, Queen’s University , Kingston, Ontario K7L 3N6 , Canada
2. Department of Biology, University of New Brunswick , Fredericton, New Brunswick E3B 5A3 , Canada
3. Atlantic Forestry Centre , Natural Resources Canada– , 1350 Regent St, Fredericton, New Brunswick E3B 5P7 , Canada
4. Canadian Forest Service , Natural Resources Canada– , 1350 Regent St, Fredericton, New Brunswick E3B 5P7 , Canada