1. School of Life Sciences and Biotechnology, The Joint International Research Laboratory of Metabolic and Developmental Sciences, Shanghai Jiao Tong University , Shanghai 200240 , China
2. Shanghai Collaborative Innovation Center of Agri-Seeds/Joint Center for Single Cell Biology, Shanghai Jiao Tong University , Shanghai 200240 , China
3. School of Agriculture and Biology, Joint Center for Single Cell Biology, Shanghai Jiao Tong University , Shanghai 200240 , China
4. State Key Laboratory of Genetic Engineering and Ministry of Education Key Laboratory of Biodiversity Science and Ecological Engineering and Institute of Biodiversity Sciences, Institute of Plant Biology, Center for Evolutionary Biology, School of Life Sciences, Fudan University , Shanghai 200240 , China