1. College of Agriculture and Biotechnology, Zhejiang University , Zijingang Campus, Hangzhou 310058 , China
2. School of Horticulture, Anhui Agricultural University , Hefei 230036 , China
3. The New Zealand Institute for Plant & Food Research Limited (Plant & Food Research) , Mt Albert, Private Bag 92169, Auckland Mail Centre, Auckland 1142 , New Zealand
4. State Key Laboratory of Plant Physiology and Biochemistry, College of Life Science, Zhejiang University , Zijingang Campus, Hangzhou 310058 , China
5. School of Biological Sciences, University of Auckland , Private Bag 92019, Auckland 1010 , New Zealand