1. Institute of Nuclear Agricultural Sciences, Key Laboratory of Nuclear Agricultural Sciences of Ministry of Agriculture and Zhejiang Province, College of Agriculture and Biotechnology, Zhejiang University , Hangzhou 310058 , China
2. Weill Institute for Cell and Molecular Biology and School of Integrative Plant Science, Section of Plant Biology, Cornell University , Ithaca, NY 14853 , USA
3. Section of Cell and Developmental Biology, Division of Biological Sciences, University of California San Diego , La Jolla, CA 92093 , USA