1. National Key Laboratory of Plant Molecular Genetics, CAS Center for Excellence in Molecular Plant Sciences, Shanghai Institute of Plant Physiology and Ecology, Chinese Academy of Sciences , Shanghai 200032 , China
2. University of the Chinese Academy of Sciences , Beijing 100049 , China
3. Institute for Agricultural Biosciences, Oklahoma State University , 3210 Sam Noble Parkway, Ardmore, Oklahoma 73401 , USA