1. College of Agriculture and Biotechnology, Zhejiang University, Zijingang Campus , Hangzhou 310058 , China
2. Zhejiang Provincial Key Laboratory of Horticultural Plant Integrative Biology, Zhejiang University, Zijingang Campus , Hangzhou 310058 , China
3. The State Agriculture Ministry Laboratory of Horticultural Plant Growth, Development and Quality Improvement, Zhejiang University, Zijingang Campus , Hangzhou 310058 , China
4. Plant Biology Section, School of Integrative Plant Science, Cornell University , Ithaca, NY 14853 , USA
5. Division of Plant and Crop Sciences, School of Biosciences, University of Nottingham, Sutton Bonington Campus , Loughborough LE12 5RD , UK
6. United States Department of Agriculture – Agricultural Research Service and Boyce Thompson Institute for Plant Research, Cornell University , Ithaca, NY 14853 , USA