1. The Sainsbury Laboratory, University of East Anglia , Norwich Research Park, Norwich NR4 7UH , UK
2. Laboratory of Crop Evolution, Graduate School of Agriculture, Kyoto University , Mozume, Muko, Kyoto 617-0001 , Japan
3. PRESTO, Japan Science and Technology Agency , 4-1-8, Honcho, Kawaguchi, Saitama 332-0012 , Japan
4. Agronomy, Horticulture and Plant Sciences Department, South Dakota State University , Brookings, SD 57007 , USA
5. Plant Genomic Network Research Team, RIKEN Center for Sustainable Resource Science , 1-7-22 Suehiro-cho, Tsurumi-ku, Yokohama, Kanagawa 230-0045 , Japan
6. Plant Epigenome Regulation Laboratory, RIKEN Cluster for Pioneering Research , 2-1 Hirosawa, Wako, Saitama 351-0198 , Japan
7. Kihara Institute for Biological Research, Yokohama City University , 641-12 Maioka-cho, Totsuka-ku, Yokohama, Kanagawa 244-0813 , Japan