1. Department of Biology, University of Massachusetts , Amherst, MA 01003 , USA
2. Department of Botany and Plant Sciences, University of California , Riverside, CA 92521 , USA
3. Department of Botany, University of New Mexico , Albuquerque, NM 87131 , USA
4. Division of Biological Sciences, University of California , Riverside, CA 92093 , USA
5. Department of Biological Sciences, University of North Texas , Denton, TX 76203 , USA