1. Key Laboratory of Tobacco Gene Resources, Biotechnology Center, Tobacco Research Institute, Chinese Academy of Agricultural Sciences (CAAS) , Qingdao 266101 , China
2. Graduate School of Chinese Academy of Agricultural Science , Beijing 100081 , China
3. Laboratory of Biomass and Green Technologies, Gembloux Agro-Bio Tech, University of Liege , Gembloux 5030 , Belgium
4. Technology Center, China Tobacco Jiangsu Industrial Co., Ltd. , Nanjing 210019 , China
5. Key Laboratory of Natural Products Synthetic Biology of Ethnic Medicinal Endophytes, State Ethnic Affairs Commission, Yunnan Minzu University , Kunming 650031 , China
6. College of Agronomy, Qingdao Agricultural University , Qingdao 266109 , China
7. Key Laboratory of Biofuels, Qingdao Engineering Research Center of Biomass Resources and Environment, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences , Qingdao 266101 , China