1. Evolutionary Biology Centre, Department of Population Biology, Uppsala University, Norbyvägen 18 D,SE-75236 Uppsala, Sweden
2. Departamento de Fisiología y Biología Animal, Universidad de Sevilla, Av. Reina Mercedes 6, 41071 Sevilla, Spain;
3. Dipartimento di Biologia Animale, Università di Pavia, Piazza Botta 9, 27100 Pavia, Italy