1. Macroevolution and Macroecology, Research School of Biology, Australian National University, Canberra, ACT 0200, Australia
2. Centre of Excellence for the Dynamics of Language, Australian National University, Canberra, ACT 0200, Australia
3. Mathematical Sciences Institute, Australian National University, Canberra, ACT 0200, Australia
4. Karungkarni Arts, via Katherine, Kalkaringi, NT 0852, Australia
5. School of Languages and Cultures, University of Queensland, Brisbane, QLD 4072, Australia