1. Key Laboratory of Ecosystem Network Observation and Modelling, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences , Beijing 100101 , China
2. College of Resources and Environment, University of Chinese Academy of Sciences , Beijing 100190 , China
3. State Key Laboratory of Tibetan Plateau Earth System, Resources and Environment (TPESRE), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing 100101 , China