1. INRAE, Bordeaux Sciences Agro, UMR 1391 ISPA , F-33882 Villenave-d’Ornon , France
2. CIRAD, PERSYST Department, UPR 115 AIDA , 97490 Sainte-Clotilde, La Reunion , France
3. AIDA, PERSYST Department, Univ Montpellier, CIRAD , Montpellier , France
4. New Zealand Forest Research Institute trading as Scion , 8440 Christchurch , New Zealand
5. Department of Botany, Ecology and Plant Physiology, University of La Laguna , 38206 La Laguna , Spain